Georgia Troup’s 2021 thesis is entitled
Understanding the influence of nutritional drivers on the habitat use of African elephants (Loxodonta africana) living in a semi-arid, anthropogenic landscape.

She concludes that her ‘Results also highlight the success of Rukinga in providing a safe refuge for elephants when travelling outside the national parks and into areas of human disturbance. Ultimately, the findings of this thesis highlight how the habitat use of elephants in Tsavo is influenced by nutritional drivers, specifically protein.’
For more details see
https://library.anu.edu.au/search~S1?/Ygeorgia+troup&Submit=GO&SORT=D/Ygeorgia+troup&Submit=GO&SORT=D&search=georgia+troup&SUBKEY=georgia+troup/1%2C2%2C2%2CB/frameset&FF=Ygeorgia+troup&Submit=GO&SORT=D&1%2C1%2C